Finally，for the many small mammals that supplement their insect diet with fruits or seeds an inability to span open gaps between tree crowns may be problematic，since trees that yield these foods can be sparse.
(特殊结构supplement…with…——以…补充)
最后，对于以水果或种子来补充其昆虫食物的许多小型哺乳动物而言，不能跨越树冠之间的无树问隙可能很成问题，因为生产这些食物的树林可能分布很稀疏。
分句1：for the many small mammals
分句2：that supplement their insect diet with fruits or seeds an inability to span open gaps between tree crowns may be problematic
分句3：since trees that yield these foods can be sparse
1和2嵌套
1和3并列
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