Missing until recently were fossils clearly intermediate，or transitional，between land mammals and cetaceans.
(倒装句)
到最近为止，一直缺乏连接陆地哺乳动物和鲸目动物的明显的中间或过渡化石。
分句1：Missing until recently were fossils clearly intermediate
分句2：or transitional
分句3：between land mammals and cetaceans
1和2并列
1、2和3嵌套
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