Footprints thus provide US not merely with rare impressions of the soft tissue of early hominids，but also with evidence of upright walking that in many ways is clearer than can be obtained from the analysis of bones.
(特殊结构not merely…but also;比较结构)
因此脚印不仅仅为我们提供早期人类软组织的稀有印记，而且为我们提供了直立行走的证据，这些证据在许多方面比骨头分析所提供的证据更清楚。
分句1. Footprints thus provide US not merely with rare impressions of the soft tissue of early hominids
分句2. but also with evidence of upright walking
分句3. that in many ways is clearer than can be obtained from the analysis of bones
1和2 并列
2和3嵌套
2中省略：Footprints provide;3是2的同位语从句
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